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The Appendages, Anatomy, and Relationships of Trilobites

TABLE OF CONTENTS.

Reading edition

Historical review 17

Part I. The appendages of trilobites 20

Terminology 20

The appendages of Neolenus 21

Historical 21

Neolenus serratus (Rominger) 21

Cephalon 21

Thorax 22

Pygidium 23

Epipodites and exites 23

Description of individual specimens 23

Restoration of Neolenus 30

Nathorstia transitans Walcott 31

The appendages of Isotelus 32

Historical 32

Isotelus latus Raymond 34

Isotelus maximus Locke 35

Restoration of Isotelus 37

Isotelus gigas Dekay 37

Isotelus arenicola Raymond 39

The appendages of Triarthrus (see also Part IV) 39

Triarthrus becki Green 39

Historical 40

Restoration of Triarthrus 42

Relation of cephalic appendages to marking on dorsal surface of glabella 43

Anal plate 44

The appendages of Ptychoparia 45

Ptychoparia striata (Emmrich) 45

Ptychoparia cordilleræ (Rominger) 45

Ptychoparia permulta Walcott 45

The appendages of Kootenia 46

Kootenia dawsoni Walcott 46

The appendages of Calymene and Ceraurus 46

Historical 46

Comparison of the appendages of Calymene and Ceraurus with those of Triarthrus 47

Spiral branchiæ 48

Ventral membrane 50

Appendifers 51

Calymene senaria Conrad 52

Cephalic appendages 52

Thoracic appendages 53

Pygidial appendages 54

Relation of hypostoma to cephalon in Calymene 55[12]

Restoration of Calymene 56

Calymene sp. ind. 56

Ceraurus pleurexanthemus Green 57

Cephalic appendages 58

Thoracic appendages 59

Pygidial appendages 59

Relation of hypostoma to cephalon 59

Restoration of Ceraurus pleurexanthemus 60

The appendages of Acidaspis trentonensis Walcott 61

The appendages of Cryptolithus (see also Part IV) 61

Cryptolithus tessellatus Green 61

Restoration of Cryptolithus 62

Summary on the ventral anatomy of trilobites 64

Comparison of appendages of different genera 64

Coxopodite 64

Cephalon 64

Thorax 66

Pygidium 67

Caudal rami 68

Homology of cephalic appendages with those of other Crustacea 69

Functions of the appendages 70

Antennules 70

Exopodites 70

Endopodites 71

Use of the pygidium in swimming 72

Coxopodites 74

Position of the appendages in life 74

Part II. Structure and habits of trilobites 77

Internal organs and muscles 77

Alimentary canal 77

Ceraurus pleurexanthemus 79

Calymene senaria 80

Cryptolithus goldfussi 80

Summary 81

Gastric glands 82

Summary 84

Heart 85

Illænus 85

Ceraurus and Calymene 85

The median "ocellus" or "dorsal organ" 86

Nervous system 89

Various glands 89

Dermal glands 89

Renal excretory organs 90

Reproductive organs 90

Panderian organs 90[13]

Musculature 91

Flexor muscles 92

Extensor muscles 92

Hypostomial muscles 94

Eyes 96

Summary 97

Sex 98

Eggs 98

Methods of life (See also under "Functions of the Appendages") 98

Habits of locomotion 99

Food and feeding methods 103

Tracks and trails 104

Part III. Relationship of the trilobites to other Arthropoda 106

Crustacea 106

Branchiopoda 106

Burgessia bella Walcott 108

Waptia fieldensis Walcott 108

Yohoia tenuis Walcott 109

Opabina regalis Walcott 109

Summary 109

Copepoda 110

Archicopepoda 111

Ostracoda 112

Cirripedia 113

Malacostraca 113

Phyllocarida 113

Syncarida 114

Isopoda 114

Marrella splendens Walcott 115

Restoration of Marrella 116

Arachnida 117

Trilobites not Arachnida 117

Merostomata 119

Sidneyia inexpectans Walcott 119

Emeraldella brocki Walcott 119

Molaria and Habelia 120

Araneæ 121

Insecta 122

Chilopoda 123

Diplopoda 124

Primitive characteristics of trilobites 125

Trilobites the most primitive arthropods 125

Limbs of trilobites primitive 125

Summary 128

Number of segments in the trunk 128

Form of the simplest protaspis 132[14]

Origin of the pygidium 134

Width of the axial lobe 137

Presence or absence of a "brim" 137

Segmentation of the glabella 137

Summary 138

The simplest trilobite 138

Naraoia compacta Walcott 139

The ancestor of the trilobites, and the descent of the Arthropoda 140

Evolution within the Crustacea 142

Summary 144

Evolution of the Merostomata 146

Evolution of the "Tracheata" 147

Summary on lines of descent 147

Final summary 151

Part IV. Description of the appendages of individual specimens 152

Triarthrus becki Green 152

Cryptolithus tessellatus Green 158

Bibliography 163

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