SYSTEMATIC CATALOGUE

OF THE

EOCENE VERTEBRATES OF WYOMING.

Although we have endeavored to make this catalogue as complete and accurate as possible, we feel that errors and omissions are, from the nature of the case, unavoidable. The material is so vast and heterogeneous, and is so much scattered throughout numberless reports, bulletins, journals, and the like, that its correct compilation is attended with great difficulties. However, a beginning is here made which will render subsequent work easier. Owing to the great number of discoveries which have since been made, Dr. Leidy's excellent catalogue of 1871 covers now but a very small portion of the ground.

In this list we have made no attempt to decide disputed questions of priority and synonymy. Doubtless, many of the genera and species here given will be found to be synonyms of American or European forms; but at present these questions cannot be settled. Where synonyms are given, they are the determinations of the original describer indicated in subsequent publications

MAMMALIA.

PRIMATES.

Notharctus, Leidy. Pr. Ac. Nat. Sc., 1870, p. 114; Cont. to Ext. Vert. Faun., p. 86; Ann. Rep. U. S. Geol. Survey of Terrs., 1871, p. 364. N. tenebrosus, Leidy, loc. cit.

Hyopsodus, Leidy. Pr. Ac. Nat. Sc., 1870, p. 110; U. S. Survey of Terrs., 1870, p. 354; do. 1871, p. 362; Cont. to Ext. Vert. Faun., p. 75.

H. paulus, Leidy, loc. cit.

H. minusculus, Leidy. Cont. to Ext. Vert. Faun., p. 81.

H. gracilis, Marsh. Am. Journ. of Sci., vol. ii., p. 42.

H. vicarius, Cope. Described in U. S. Survey of Terrs., 1872, p. 609, as Microsyops vicarius. Wheeler's Survey, vol. iv., pt. ii., p. 150.

Microsyops, Leidy. Pr. Ac. Nat. Sc., 1872, p. 20; U. S. Survey of Terrs., 1871, p. 363; Cont. to Ext. Vert. Faun., p. 82.

M. gracilis, Leidy, loc. cit. In the "Contributions," Dr. Leidy, considering Limnotherium, Marsh, as a synonym of Microsyops, has called this species M. elegans.

Hipposyus, Leidy. Pr. Ac. Nat. Sc., 1872, p. 37; Cont. to Ext. Vert. Faun., p. 90.

H. formosus, Leidy, loc. cit.

H. robustior, Leidy Cont. to Ext. Vert. Faun., p. 93; Notharctus robustior. U. S. Geol. Survey of Terrs., 1871, p. 364.

Antiacodon, Marsh. Am. Journ. of Sci., 3d Ser. vol. iv., p. 210.

A. venustus, Marsh, loc. cit.

Bathrodon, Marsh. Am. Journ. of Sci., 3d Ser. vol. iv., p. 211.

B. typus, Marsh, loc. cit.

Lemuravus, Marsh. Am. Journ. of Sci., 3d Ser. vol. ix., p. 239.

L. distans, Marsh, loc. cit.

Limnotherium, Marsh. Am. Journ. of Sci., 3d Ser. vol. ii., p. 43.

L. tyrannus, Marsh, loc. cit.

L. elegans, Marsh, loc. cit.

L. affine, Marsh. Am Journ. of Sci., 3d Ser. vol. iv., p. 207.

Mesacodon, Marsh. Am. Journ. of Sci., 3d Ser. vol. iv., p. 212.

M. speciosus, Marsh, loc. cit.

Palæacodon, Leidy. Pr. Ac. Nat. Sc., 1872; p. 21. U. S. Geol. Survey of Terrs., 1871, p. 356; Cont. to Ext. Vert. Faun., p. 122.

P. verus, Leidy, loc. cit.

P. vagus, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 224.

Telmatolestes, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 206.

T. crassus, Marsh, loc. cit.

Tomitherium, Cope. Pr. Am. Phil. Soc., 1872, p. 470; U. S. Geol. Survey of Terrs., 1872, p. 546; Wheeler's Survey, vol. iv., p. 135, pt. ii.

T. rostratum, Cope, loc. cit.; U. S. Survey, 1872, p. 548.

Anaptomorphus, Cope. Pr. Am. Phil. Soc., 1872, p. 554; U. S. Geol. Survey of Terrs., 1872, p. 549.

A. æmulus, Cope, loc. cit.

Oligotomus, Cope. On some Eocene mammals, p. 2; U. S. Geol. Survey of Terrs., 1872, p. 607.

O. cinctus, Cope, loc. cit.

Opisthotomus, Cope. Wheeler's Survey, vol. iv., p. 151, pt. ii.

O. astutus, loc. cit., p. 152.

Pantolestes, Cope. Pr. Am. Phil. Soc., 1872, p. 467; Wheeler's Survey, v. iv., pt. ii., p. 145.

P. longicaudus. Pr. Am. Phil. Soc., 1872, p. 467; Notharctus longicaudus, U. S. Geol. Survey of Terrs., 1872, p. 549.

Sarcolemur, Cope. Pr. Ac. Nat. Sc., 1875, p. 256; Wheeler's Survey, v. iv., pt. ii., p. 147.

S. pygmæus, Cope. Lophiotherium pygmæus, Pr. Am. Phil. Soc., 1872, extras July 20; Antiacodon pygmæus, U. S. Geol. Survey of Terrs., 1872, p. 607; Hyopsodus pygmæus, Pr. Am. Phil. Soc., 1872, p. 461.

S. furcatus, Cope. Antiacodon furcatus. On some Eocene mammals, p. i.; U. S. Geol. Survey of Terrs., 1872, p. 608.

Thinolestes, Marsh. Am. Journ. Sc., 3d Ser. vol. iv., p. 205.

T. anceps, Marsh, loc. cit.

CARNIVORA.

Uintacyon, Leidy. Pr. Ac. Nat. Sc., 1872, p. 277; Cont. to Ext. Vert. Faun., p. 118.

U. edax, Leidy, loc. cit.

U. vorax, Leidy, loc. cit. Cont. to Ext. Vert. Faun., p. 120.

Sinopa, Leidy. Pr. Ac. Nat. Sc., 1871, p. 115; U. S. Geol. Survey of Terrs., 1871, p. 355; Cont. to Ext. Vert. Faun., p. 116.

S. rapax, Leidy, loc. cit.

S. eximia, Leidy. Cont. to Ext. Vert. Faun., p. 118.

Canis.

C. montanus, Marsh. Am. Jour. of Sc., 3d Ser. vol. ii., p. 123.

Dromocyon, Marsh. Am. Jour. of Sc., 3d Ser. vol. xii., p. 403.

D. vorax, Marsh, loc. cit.

Harpalodon, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 216.

H. sylvestris, Marsh, loc cit.

H. vulpinus, Marsh, loc. cit., p. 217.

Limnocyon, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 126.

L. verus, Marsh, loc. cit.

L. riparius, Marsh, loc. cit., p. 203.

L. agilis, Marsh, loc. cit., p. 204.

Limnofelis, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 202.

L. ferox, Marsh, loc. cit.

L. latidens, Marsh, loc. cit., p. 203.

Oreocyon, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 406

O. latidens, Marsh, loc. cit.

Thinocyon, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 204.

T. velox, Marsh, loc. cit.

Viverravus, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 127.

V. gracilis, Marsh, loc. cit.

V. (?) nitidus, Marsh, loc. cit., p. 205.

Ziphacodon, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 216.

Z. rugatus, Marsh, loc. cit.

Vulpavus, Marsh. Am. Jour. of Sc., 3d Ser. vol. ii., p. 124.

V. palustris, Marsh, loc. cit.

Miacis, Cope. Pr. Am. Phil. Soc., 1872, p. 470.

M. parvivorus, loc. cit.; Viverravus parvivorus, U. S. Geol. Survey of Terrs., 1872, p. 560.

Mesonyx, Cope. Pr. Am. Phil. Soc., 1872. p. 460; U S. Geol. Survey of Terrs., 1872, p. 550.

M. obtusidens, Cope, loc. cit. U. S. Survey, p. 552.

Synoplotherium, Cope. Pr. Am. Phil. Soc., 1872, p. 483; U. S. Geol. Survey of Terrs., 1872, p. 554.

S. lanius, Cope, loc. cit. U. S. Survey, p. 557.

Stypolophus, Cope. Pr. Am. Phil. Soc., 1872, p. 466; U. S. Geol. Survey of Terrs., 1872, p. 559; Wheeler's Survey, vol. iv., p. 109.

S. insectivorus, Cope. Pr, Phil. Soc., 1872, p. 469; U. S. Survey, 1872, p. 557.

S. pungens, Cope, loc. cit., pp. 466, 559

S. brevicalcaratus, Cope, loc. cit., pp. 469, 560.

Megencephalon, gen. nov. This report, p 20.

M. primævus, sp. nov. This report, p. 20.

UNGULATA.

PERISSODACTYLA.

Anchitherium.

A. (?)——. This report, p. 23.

Orohippus, Marsh. Am. Jour. of Sc., vol. iv., p. 207; vol. vii., p. 247.

O. pumilus, Marsh, loc. cit., 3d Ser. vol. iv., p. 207.

O. agilis, Marsh, loc. cit., 3d Ser. vol. v., p. 407.

O. major, Marsh, loc. cit., vol. 3d Ser. vii., p. 248.

O. gracilis, Marsh, loc. cit., 3d Ser. vol. vii., p. 249; Anchitherium gracile 3d Ser. vol. ii., p. 38.

O. procyoninus, Cope. Pr. Am. Phil. Soc., 1872, p. 466; U. S. Geol. Survey, 1872, p. 606.

Palæosyops, Leidy. Pr. Ac. Nat. Sc., 1870, p. 113; 1871, pp. 114, 118, 197, 229; 1872, pp. 168, 241; U. S. Geol. Survey, 1870, p. 355; 1871, p. 358; Com. to Ext. Vert. Faun., p. 27; Cope, U. S. Geol. Survey of Terrs., 1872, p. 591.

P. paludosus, Leidy, loc. cit. Cont. to Ext. Vert. Faun., p. 28.

P. major, Leidy. U. S. Geol. Survey of Montana, 1871, p. 359; Pr. Ac. Nat. Sc., 1872, pp. 168, 241; Cont. to Ext. Vert. Faun., p. 45.

P. humilis, Leidy. Pr. Ac. Nat. Sc., 1872, p. 168, 277; Cont. to Ext. Vert. Faun., p. 58.

P. junius, Leidy. Pr. Ac. Nat. Sc., 1872, p. 277; Cont. to Ext. Vert. Faun., p. 57.

P. lævidens, Cope. Pr. Am. Phil. Soc., 1873; U. S. Survey, 1872, p. 591.

P. vallidens, Cope. Pr. Am. Phil. Soc., 1872, p. 487; loc. cit., p. 592.

P. minor, Marsh. Am. Jour. of Sc., vol. ii., p. 36.

Limnohyus, Leidy. Pr. Ac. Nat. Sc., 1872, p. 242; Cont. to Ext. Vert. Faun., p. 58; Marsh, Am. Jour. of Sc., 1872, 3d Ser. vol. iv., p. 124.

L. laticeps, Marsh. Palæosyops laticeps. Am. Journ. Sc. 3d Ser. vol. iv., p. 122.

L. robustus, Marsh, loc. cit., p. 124, is same as P. major, Leidy.

L. diaconus, Cope. U. S. Geol. Survey of Terrs., 1872, p. 593.

L. fontinalis, Cope, loc. cit., p. 594.

Telmatherium, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 123.

T. validum, Marsh, loc. cit.

Leurocephalus, gen. nov. This report, p. 42.

L. cultridens, sp. nov. This report, p. 42.

Hyrachyus, Leidy. Pr. Ac. Nat. Sc., 1871, p. 229; 1872, pp. 19, 163; U. S. Geol. Survey, 1871, p. 360; Cont. to Ext. Vert. Faun., p. 60.

H. agrarius, Leidy, loc. cit.; H. agrestis. U. S. Survey, 1871, p. 357.

H. eximius, Leidy. Pr. Ac. Nat. Sc., 1871, p. 229; 1872, p. 163; U. S. Geol. Survey of Terrs., 1871, p. 361; Cont. to Ext. Vert. Faun., p. 66; Cope, U. S. Geol. Survey of Terrs., 1872, p. 595.

H. modestus, Leidy. Pr. Ac. Nat. Sc., 1872, p. 20; U. S. Geol. Survey of Terrs., 1871, p. 361; Cont. to Ext. Vert. Faun., p. 67; Lophiodon modestus, Pr. Ac. Nat. Sc., 1870. p. 109.

H. nanus, Leidy. Pr. Ac. Nat. Sc., 3d Ser. 1872, p. 20; U. S. Survey, 1871, p. 361; Cont. to Ext. Vert. Faun., p. 67.

H. princeps, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 125.

H. implicatus, Cope. On Some Eocene Mammals, p. 5; U. S. Survey, 1872, p. 604.

H. crassidens, sp. nov. This report, p. 52.

H. imperialis, sp. nov. This report, p. 50.

H. intermedius, sp. nov. This report, p. 51.

H. paradoxus, sp. nov. This report, p. 53.

Helaletes, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 218.

H. boöps, Marsh, loc. cit.

H. latidens, sp. nov. This report, p. 54.

Lophiodon.

L bairdianus, Marsh. Am. Jour. of Sc., 3d Ser. vol. ii., p. 36.

L. affinis, loc. cit., p. 37.

L. nanus, loc. cit.

L. pumilus, loc. cit., p. 38.

Lophiotherium.

L. sylvaticum, Leidy. Pr. Ac. Nat. Sc., 1870, p. 126; Cont. to Ext. Vert. Faun., p. 69.

L. Ballardi, Marsh. Am. Jour. of Sc., 3d Ser. vol. ii., p. 39.

Orotherium, Marsh. Am: Jour. of Sc., 3d Ser. vol. iv., p. 217.

O. mintanum, Marsh, loc. cit.

Helohyus, Marsh. Am. Journ. of Sc., 3d Ser. vol. iv., p. 207.

H. plicodon, Marsh, loc. cit.

Thinotherium, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 208.

T. validum, Marsh, loc. cit.

ARTIODACTYLA.

All the forms described under this head are of uncertain reference.

Elotherium.

E. lentum, Marsh. Am. Jour. of Sc., 3d Ser. vol. ii., p. 39.

Platygonus.

P. Ziegleri, Marsh, loc cit., p. 40.

Parahyus, Marsh. Am. Jour. of Sc., 3d Ser. vol. xii., 402.

P. vagus, Marsh, loc. cit.

Homacodon, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 126.

H. vagans, Marsh, loc. cit.

Ithygrammodon, gen. nov. This report, p. 56.

I. cameloides, sp. nov. This report, p. 57.

AMBLYPODA.

Cope. Wheeler's Survey, vol., iv., pt. ii., p. 179.

DINOCERATA.

Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 344; Ibid. vol. v., pp. 117-122, 293; Ibid. vol. vi., p. 300; Ibid. vol. xi., p. 163.

Uintatherium, Leidy. Pr. Ac. Nat. Sc., 1872, p. 169; Am. Jour. of Sc., 3d Ser. vol. iv., p. 239; Cont. to Ext. Vert. Faun., p. 96.

U. robustum, Leidy, loc. cit.; Uintamastix atrox, loc. cit.

U. Leidianum, sp. nov. This report, p. 3.

U. princeps, sp. nov. This report, p. 81.

Tinoceras, Marsh. Titanotherium (?), Am. Jour. of Sc., 3d Ser. vol. ii., p. 35; Mastodon, loc. cit., vol. iv., p. 123, footnote; Tinoceras, loc. cit., vol. iv., pp. 322 and 323.

T. anceps, Marsh, loc. cit., vol. iv., p. 322.

T. grandis, Marsh, loc. cit., p. 323.

Dinoceras, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 344; Ibid. vol. v., pp. 117-122; Ibid. vol. v., p. 408.

D. mirabilis, Marsh, loc. cit., vol. iv., p. 344.

D. lucaris, Marsh, loc. cit., vol. v., p. 408.

D. lacustre, Marsh, loc. cit., vol. iv., p. 344.

Loxolophodon, Cope. Pr. Am. Phil. Soc., 1872, pp. 580, 488; U. S. Survey, 1872, p. 565.

L. cornutus, Cope, loc. cit.; U. S. Survey, 1872, p. 568.

Eobasileus, Cope. Pr. Am. Phil. Soc., 1872, p. 485; U. S. Survey, 1872, p. 575.

E. pressicornis, Cope. Pr. Am. Phil. Soc., 1872, p. 580; loc. cit.

E. furcatus, Cope. U. S. Survey, 1872, p. 580; Loxolophodon furcatus, Pr. Am. Phil. Soc., 1872, p. 580.

Coryphodon.

C. hamatus, Marsh. Am. Jour. of Sc., 3d Ser. vol. xi., p. 425.

Bathmodon, Cope. Pr. Am. Phil. Soc., 1872, p. 417; U. S. Geol. Survey, 1871, p. 350; 1872, p. 586.

B. radians, Cope, loc. cit.

B. semicinctus, Cope, loc. cit.

B. latipes, Cope, loc. cit.

Metalophodon, Cope. Pr. Am. Phil. Soc., 1872, p. 542; U. S. Geol. Survey, 1872, p. 589.

M. armatus, Cope, loc. cit.

TILLODONTIA.

Marsh. Am. Jour. of Sc., 3d Ser. vol. ix., p. 221; vol. xi., p. 249.

Anchippodus, Leidy. Pr. Ac. Nat. Soc., 1868, p. 232; Ext. Mam. N. Am., p. 403.

Trogosus. Pr. Ac, 1871, p. 113; Cont. to Ext. Vert. Faun., p. 71.

A. riparius, Leidy. Pr. Ac. Nat. Soc., 1868, p. 232; Cont. to Ext. Vert. Faun., p. 71 (as Trogosus castoridens).

A. vetulus, Leidy; Trogosus vetulus, Pr. Ac. Nat. Sc., 1871, p. 229; Cont. to Ext. Vert. Faun., p. 75.

Tillotherium. Am. Jour. of Sc., 3d Ser. vol. v., p. 485; Ibid. vol. xi., p. 249.

T. hydracoides, Marsh, loc. cit.

T. latidens, Marsh, loc. cit., vol. vii., p. 533.

T. fodiens, Marsh, loc. cit., vol. ix., p. 241.

Stylinodon, Marsh. Am. Jour. of Sc., 3d Ser. vol. vii., p. 532.

S. mirus, loc. cit.

RODENTIA.

Paramys, Leidy. Pr. Ac. Nat. Sc., 1871, p. 231; U. S. Geol. Survey, 1871, p. 357; Cont. to Ext. Vert. Faun., p. 110.

P. delicatus, Leidy, loc. cit.

P. delicatior, Leidy, loc. cit.

P. delicatissimus, Leidy, loc. cit.; Cont. to Ext. Vert. Faun., p. 111.

P. robustus, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 218.

P. superbus, sp. nov. This report, p. 84.

P. leptodus, Cope. U. S. Geol. Survey, 1872, p. 609.

Mysops, Leidy. Pr. Ac. Nat. Sc., 1871, p. 232; U. S. Geol. Survey, 1871, p. 357; Cont. to Ext. Vert. Faun., p. 111.

M. minimus, Leidy, loc. cit.

M. fraternus, Leidy. Cont. to Ext. Vert. Faun., p. 112.

Pseudotomus, Cope. Pr. Am. Phil. Soc., 1872, p. 467; U. S. Geol. Survey, 1872, p. 610.

P. hians, Cope, loc. cit.

Arctomys, Marsh. Am. Jour. of Sc., 3d Ser. vol. ii., p. 121.

A. vetus, Marsh, loc. cit.

Geomys, Marsh. Am. Jour. of Sc., 3d Ser. vol. ii., p. 121.

G. bisulcatus, Marsh, loc. cit.

Sciuravus, Marsh. Am. Jour. of Sc., 3d Ser. vol. ii., p. 122.

S. nitidus, Marsh, loc. cit.

S. undans, Marsh, loc. cit.

S. parvidens, Marsh, loc. cit., 3d Ser. vol. iv., p. 220.

Tillomys, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 219.

T. senex, Marsh, loc. cit.

T. parvus, Marsh, loc. cit.

Tachymys, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 219.

T. lucaris, Marsh, loc. cit.

Colonymys, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 220.

C. celer, Marsh, loc. cit.

INSECTIVORA.

Omomys, Leidy. Pr. Ac. Nat. Sc., 1869, p. 63; Ext. Mam. of N. Am., p. 408; Cont. to Ext. Vert. Faun., p. 120.

O. Carteri, Leidy, loc. cit.

Washakius, Leidy. Cont. to. Ext. Vert. Faun., p. 123.

W. insignis, Leidy, loc. cit.

Passalacodon, Marsh. Am. Jour. of Sc., vol. iv. p. 208.

P. litoralis, Marsh, loc. cit.

Anisacodon,[1] Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 209.

A. elegans, Marsh, loc. cit.

Centetodon, Marsh, Am. Jour. of Sc., 3d Ser. vol. iv., p. 209.

C. pulcher, Marsh, loc. cit.

C. altidens, Marsh, loc. cit., p. 214.

Hemiacodon, Marsh, Am. Jour. of Sc., 3d Ser. vol. iv., p. 212.

H. gracilis, Marsh, loc. cit.

H. nanus, Marsh, loc. cit., p. 113

H. pucillus, Marsh, loc. cit.

Entomodon, Marsh, Am. Jour. of Sc., 3d Ser. vol. iv., p. 214.

E. comptus, Marsh, loc. cit.

Entomacodon, Marsh, Am. Jour. of Sc., 3d Ser. vol. iv., p. 214.

E. minutus, Marsh, loc. cit.

E. angustidens, Marsh, loc. cit., p. 222.

Apatemys, Marsh, Am. Journ. of Sc., 3d Ser. vol. iv., p. 221.

A. bellus, Marsh, loc. cit.

A. bellulus, Marsh, loc. cit.

Talpavus, Marsh, Am. Journ. of Sc., 3d Ser. vol. iv., p. 128.

T. nitidus, Marsh, loc cit.

CHIROPTERA.

Nyctitherium, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 127.

N. velox, Marsh, loc. cit.

N. priscum, Marsh, loc. cit., p. 128.

Nyctilestes, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 215.

N. serotinus, Marsh, loc. cit.

MARSUPIALIA.

Triacodon, Marsh. Am. Jour. of Sc., 3d Ser. vol. ii., p. 123.

T. fallax, Marsh, loc. cit.

T. grandis, Marsh, loc. cit. 3d Ser. vol. iv., p. 222.

T. nanus, Marsh, loc. cit., p. 223.

T. aculeatus, Cope. Pr. Phil. Soc., 1872, p. 460; U. S. Geol. of Terrs., 1872, p. 611.

Genera incertæ sedis.

Stenacodon, Marsh. Am. Jour. of Sc., 3d Ser. vol. iv., p. 210.

S. rarus, Marsh, loc. cit.