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Alpine flora of the Canadian Rocky Mountains

Front matter

Reading edition

Mimulus Lewisii Pursh. (⅔ Nat.) Red Monkey-Flower.

Alpine Flora of the Canadian Rocky Mountains

By Stewardson Brown

Curator of Herbarium Academy of Natural Sciences Philadelphia

Illustrated with Water-Colour Drawings and Photographs

By Mrs. Charles Schäffer

G. P. Putnam’s Sons New York and London The Knickerbocker Press 1907

Copyright, 1907 BY G. P. PUTNAM’S SONS

OFFERED TO THE LOVERS OF ALPINE FLORÆ IN THE MEMORY OF DR. CHARLES SCHÄFFER OF PHILADELPHIA, WHO WAS> AMONG THE PIONEER BOTANISTS OF THE CANADIAN ROCKIES, AND WHO EARLY RECOGNISED THIS REGION AS A NEW AND INTERESTING FIELD FOR STUDY

PREFACE

The present volume, though prepared in manual form, is meant only as a guide to the rich and interesting flora of the Canadian Rockies and Selkirks or those portions traversed by the Canadian Pacific Railway between Banff and Glacier. While many of the plants herein described are found throughout the entire region, yet the species characteristic of the two mountain ranges are vastly different. For the most part those of the Rockies might be characterised as plants adapted to the withstanding of severe conditions of drouth and cold, being mostly low and tufted, with small surfaces of leaf exposure, either thick and leathery or in many instances with an ample covering of protecting hairs; while those of the Selkirks, owing to the more humid atmospheric conditions, are essentially moisture-loving forms, with a luxuriant growth of stems and leaves; it is only where the conditions are similar in the two regions that we find the same or similar forms existing. Few of the more characteristic Rocky Mountain species extend west of the divide, while of those of the Selkirks, few extend east of it, except where there is a luxuriant forest growth, with the consequent retention of moisture, and then only at altitudes of from 2000 to 3000 feet greater.

Contrasting the plants of this region with those of the European Alps it is interesting to note, that while the species are for the most part vastly different, yet there is a certain close resemblance in the families and genera which are represented; the Anemones are here, but not in the scarlets and crimsons, running more to whites and purples, and so are the Buttercups, dwarf many of them, but with large, showy flowers; the Saxifrages and Drabas are innumerable as are the Vetches and also the Heaths, but the latter with fewer and very different species. Some plants, however, like the White Mountain Avens (Dryas octopetala), the Butterwort (Pinguicula vulgaris), the Moss Campion (Silene acaulis), the Mountain Cranberry (Vitis-idæa) and the Low Cranberry (Oxycoccus), the One-flowered Wintergreen (Moneses uniflora), the Forget-me-not (Myosotis alpestris) and others are found on both sides of the Atlantic, yet we have no such array of Primroses or Gentians as our European brothers, though both genera are represented with us, while on the other hand they can boast of nothing comparable to our Indian Paint-Brush, found throughout the region on the river shores and bars during late June and early July and later through the summer in its innumerable forms and colours in the moist alpine meadows and slopes.

The ferns and their allies with the more striking of the trees and shrubs have been included in the present work, together with the majority of the herbaceous flowering plants, though some of the largely represented but less striking groups such as the Grasses, Sedges, and Willows have been entirely omitted, not seeming of sufficient general interest to warrant the space required for their intelligent treatment. The species are arranged in accordance with their scientific relationships, with keys to the genera and a general key to the families. Among the illustrations herein contained, many of the plants of the Northwest are figured for the first time.

Acknowledgment is here made to Messrs. David McNicoll, Robert Kerr, and other officials and employees of the Canadian Pacific Railway Company through whose courtesy was made possible the gathering of the data for the basis of this work.

S. B.

Academy of Natural Sciences, Philadelphia, October 28, 1907.

CONTENTS

PAGE

Preface v

Glossary xxv

General Key to the Families xxxiii

Adder’s-Tongue Family 1

Fern Family 3

Horsetail Family 14

Club-moss Family 18

Selaginella Family 22

Pine Family 23

Yew Family 34

Arum Family 35

Bunch-Flower Family 36

Lily Family 41

Lily-of-the-Valley Family 45

Iris Family 52

Orchid Family 53

Willow Family 68

Birch Family 70

Mistletoe Family 72

Sandalwood Family 73

Buckwheat Family 75

Purslane Family 80

Pink Family 81

Crowfoot Family 90

Barberry Family 109

Poppy Family 110

Mustard Family 111

Stone-Crop Family 125

Grass-of-Parnassus Family 126

Saxifrage Family 128

Gooseberry Family 146

Rose Family 150

Apple Family 165

Plum Family 167

Pea Family 168

Flax Family 182

Crowberry Family 183

Staff-Tree Family 184

Maple Family 185

St. John’s-wort Family 186

Violet Family 187

Oleaster Family 191

Evening Primrose Family 193

Ginseng Family 199

Celery Family 201

Dogwood Family 205

Wintergreen Family 207

Heath Family 211

Huckleberry Family 220

Primrose Family 226

Gentian Family 231

Buckbean Family 236

Dogbane Family 237

Water-Leaf Family 238

Borage Family 240

Mint Family 244

Figwort Family 248

Butterwort Family 262

Madder Family 263

Honeysuckle Family 264

Valerian Family 271

Bellflower Family 273

Chicory Family 275

Thistle Family 283

Index 323

ILLUSTRATIONS

PLATE FACING PAGE

Mimulus Lewisii Pursh. Red Monkey-Flower Frontispiece

1 Pinus albicaulis Engelm. White Pine 24

1 Pinus Murrayana Oreg. Com. Jack Pine 24

2 Larix Lyallii Parl. Lyall’s Larch 26

2 Thuja plicata Don. Giant Cedar 26

3 Abies lasiocarpa (Hook) Nutt. Balsam Fir 28

3 Pseudotsuga mucronata (Raf.) Sudw. Douglas Fir 28

4 Tsuga Mertensiana (Bong.) Carr. Mountain Hemlock 30

4 Tsuga heterophylla (Raf.) Sarg. Hemlock 30

5 Picea albertiana. S. Brown. Alberta Spruce 32

5 Picea albertiana. S. Brown. Alberta Spruce 32

6 Zygadenus elegans Pursh. Tall Zygadenus 38

6 Zygadenus gramineus Rydb. Zygadenus 38

7 Stenanthella occidentalis (A. Gray). Rydb. Stenanthium 40

7 Clintonia uniflora Kunth. Clintonia 40

8 Tofieldia intermedia Rydb. False Asphodel 42

8 Vagnera stellata (L) Morong. Star-Flowered Solomon’s Seal 42

9 Erythronium grandiflorum. Pursh. Snow Lily 44

10 Lilium montanum A. Nels. Red Mountain-Lily 46

10 Kruhsea streptopoides (Ledeb) Kearney. Kruhsea 46

11 Cypripedium passerinum Rich. Small White Lady’s Slipper 54

11 Cypripedium parviflorum Salisb Small Yellow Lady’s Slipper 54

12 Corallorhiza Corallorhiza (L). Karst. Coral-Root 56

12 Lyschiton kamtschatcense (L.) Schott. Western Skunk-Cabbage 56

13 Cytherea bulbosa (L.) House. Calypso 58

13 Orchis rotundifolia Pursh. Small Round-Leaved Orchid 58

14 Limnorchis dilatatiformis Rydb. Purplish-Green Bog-Orchid 62

14 Peramium Menziesii (Lindl.) Morong. Menzies’ Rattlesnake Plantain 62

15 Ophrys borealis (Morong). Northern Twayblade 64

15 Ophrys nephrophylla Rydb. Heart-Shaped Twayblade 64

15 Cœloglossum bracteatum (Willd) Parl. Long-Bracted Orchid 64

15 Limnorchis fragrans Rydb. Fragrant White Bog-Orchid 64

15 Lysiella obtusata (Pursh) Rydb. Small Northern Bog-Orchid 64

16 Comandra pallida DC White Comandra 74

16 Eriogonum subalpinum Greene. Tall White Eriogonum 74

17 Claytonia lanceolata Pursh. Spring Beauty 80

17 Claytonia parvifolia Moc. Small-Leaved Spring Beauty 80

18 Silene acaulis L. Moss Campion 82

18 Lychnis apetala L. Nodding Lychnis 82

19 Alsine borealis (Bigel.) Britton. Northern Stitchwort 86

19 Mœhringia lateriflora (L.) Blunt-Leaved Sandwort 86

19 Arenaria capillaris nardifolia (Ledeb.) Regel. Rock Sandwort 86

20 Caltha leptosepala Hook. Caltha leptosepala 104

20 Trollius albiflorus (A. Gray) Rydb. Western Globe-Flower 104

21 Aquilegia brevistyla Hook. Small Blue Columbine 102

22 Aquilegia flavescens S. Wats. Yellow Columbine 106

22 Aquilegia formosa Fisch. Western Columbine 106

23 Delphinium Brownii Rydb. Mountain Larkspur 108

23 Anemone globosa Nutt. Wind-Flower 108

24 Anemone Drummondii S. Wats. Alpine Anemone 92

24 Anemone parviflora Michx. Northern Anemone 92

25 Pulsatilla hirsutissima (Pursh) Britton Pasque-Flower 94

25 Pulsatilla occidentalis (S. Wats.) Freyn. Western Anemone 94

26 Atragene columbiana Nutt. Purple Virgin’s-Bower 90

27 Ranunculus saxicola Rydb. Ranunculus saxicola 100

27 Ranunculus alpeophilus. A Nels. Ranunculus alpeophilus 100

27 Ranunculus Eschscholtzii Schl. Snow Buttercup 100

28 Ranunculus eremogenes Greene Ditch Crowfoot 98

28 Ranunculus Purshii Richards. Pursh’s Buttercup 98

28 Ranunculus inamœnus Greene Ranunculus inamœnus 98

29 Thalictrum megacarpum Torr. Thalictrum megacarpum 96

29 Thalictrum occidentale A. Gray. Western Meadow Rue 96

30 Delphinium Menziesii DC. Blue Larkspur 110

30 Lithophragma parviflora (Hook.) Nutt. Lithophragma 110

31 Physaria didymocarpa (Hook.) A. Gray. Bladder-Pod 116

31 Smelowskia calycina (Desv.) C. A. Meyer. Smelowskia 116

32 Draba oligosperma Hook. Whitlow-Grass 112

32 Draba andina (Nutt.) A. Nels. Mountain Whitlow-Grass 112

33 Draba glacialis Adams Whitlow-Grass 114

33 Draba aurea Vahl. Golden Whitlow-Grass 114

34 Parnassia montanensis Rydb. & Fern Marsh Grass-of-Parnassus 126

34 Ribes lacustre (Pers.) Poir. Swamp Gooseberry 126

35 Mitella nuda L. Naked Bishop’s-Cap 132

35 Pectiantia pentandra (Hook.) Rydb. Mitrewort 132

36 Saxifraga cernua L Nodding Saxifrage 136

36 Saxifraga rivularis L. Alpine Brook Saxifrage 136

37 Micranthes Nelsoniana (D. Don.) Small. Nelson’s Saxifrage 140

37 Micranthes Lyallii (Engler) Small. Lyall’s Saxifrage 140

38 Muscaria cæspitosa (L.) Haw. Tufted Saxifrage 138

38 Micranthes rhomboidea (Greene) Small. Mountain Saxifrage 138

39 Spatularia Brunoniana (Bong.) Small. Tall Saxifrage 142

39 Leptasea austromontana (Wiegand), Small. Common Saxifrage 142

40 Tiarella unifoliata Hook. Western Foam-Flower 150

40 Lutkea pectinata (Pursh) Kuntze. Cut-Leaved Lutkea 150

41 Rubus pedatus Smith. Creeping Raspberry 152

42 Rubus parviflorus. Nutt. Salmon-Berry 156

43 Fragaria glauca (S. Wats.) Rydb. Wild Strawberry 158

43 Amelanchier florida Lindl. Service-Berry 158

44 Dryas octopetala L. White Mountain Avens 164

44 Dryas Drummondii Rich. Drummond’s Mountain Avens 164

45 Geum strictum Ait. Yellow Avens 162

45 Sieversia ciliata (Pursh) Don. Long-Plumed Purple Avens 162

46 Phaca americana (Hook.) Rydb. Arctic Vetch 172

46 Homalobus aboriginorum (Rich) Rydb. Indian Vetch 172

47 Aragallus deflexus. (Pall.) Heller Drooping Vetch 176

47 Aragallus viscidulus Rydb. Sticky Oxytrope 176

48 Aragallus Lamberti (Pursh) Greene. Loco Weed 180

48 Lathyrus ochroleucus Hook. White Vetch 180

49 Empetrum nigrum L. Black Crowberry 182

49 Linum Lewisii Pursh. Wild Blue-Flax 182

50 Pachystima myrsinites (Pursh) Raf. Mountain Lover 184

50 Viola adunca longipes (Nutt.) Rydb. Dog Violet 184

51 Elæagnus argentea Pursh. Silver-Berry 190

51 Lepargyræa canadensis (L.) Greene Buffalo-Berry 190

52 Epilobium luteum Pursh. Yellow Willow-Herb 194

52 Chamænerion latifolium (L.) Sweet. Broad-Leaved Willow-Herb 194

53 Chamænerion angustifolium (L.) Scop. Fire-Weed 196

53 Berberis aquifolium Pursh. Trailing Mahonia 196

54 Echinopanax horridum. (Smith) Dec. & Planch. Devil’s Club 200

55 Cornus canadensis intermedia Farr. Bunch-Berry 204

56 Pyrola asarifolia Michx. Liver-Leaf Wintergreen 208

56 Pyrola uliginosa Torr. Bog Wintergreen 208

56 Moneses uniflora (L.) A. Gray. One-Flowered Wintergreen 208

57 Chimaphila umbellata (L.) Nutt. Prince’s Pine 210

57 Arctostaphylos uva-ursi (L.) Spreng. Bear-Berry 210

58 Oxycoccus oxycoccus (L.) MacM. Small Cranberry 212

58 Gaultheria ovatifolia A. Gray. Ovate-Leaved Wintergreen 212

58 Vitis-idæa Vitis-idæa (L.) Britton. Mountain Cranberry 212

59 Cassiope Mertensiana (Bong.) Don. White Heath 216

59 Phyllodoce empetriformis (Smith) Don. Red False-Heather 216

59 Phyllodoce glanduliflora (Hook.) Rydb. White False-Heather 216

60 Ledum grœnlandicum Œder. Labrador Tea 218

60 Azaleastrum albiflorum (Hook.) Rydb. White Mountain Rhododendron 218

61 Menziesia ferruginea Smith. Smooth Menziesea 220

61 Kalmia microphylla (Hook.) Heller. Dwarf Swamp-Laurel 220

62 Vaccinium erythrococcum Rydb. Alpine Bilberry 222

62 Vaccinium globulare Rydb. Thin-Leaved Bilberry 222

62 Vaccinium ovalifolium Smith. Blueberry 222

63 Primula Maccalliana Wiegand. Maccalla’s Primrose 228

63 Androsace carinata Torr. Sweet Androsace 228

64 Trientalis arctica Fisch. Arctic Star-Flower 230

64 Dodecatheon conjugens Greene. Shooting-Star 230

65 Gentiana affinis Griseb. Large Gentian 234

65 Romanzoffia sitchensis Bong. Romanzoffia 234

66 Phacelia sericea (Graham) A. Gray. Mountain Phacelia 238

66 Phacelia heterophylla Pursh Phacelia heterophylla 238

67 Lappula floribunda (Lehm.) Greene. False Forget-me-not 242

67 Lithospermum linearifolium Goldie. Narrow-Leaved Puccoon 242

68 Collinsia parviflora Dougl. Small-Flowered Collinsia 248

68 Elephantella grœnlandica (Retz.) Rydb. Long-Beaked Elephantella 248

69 Pentstemon confertus Dougl. Yellow Beard-Tongue 250

69 Pentstemon pseudohumilis Rydb. Pentstemon pseudohumilis 250

70 Pentstemon fruticosus (Pursh) Greene. Large Purple Beard-Tongue 262

70 Pinguicula vulgaris L. Butterwort 262

71 Linnæa americana Forbes. American Twin-Flower 266

71 Lonicera ebractulata Rydb. Fly Honeysuckle 266

72 Lonicera involucrata (Rich.) Banks. Involucred Fly Honeysuckle 270

73 Valeriana sitchensis Bong. Wild Heliotrope 272

73 Lobelia Kalmii strictiflora Rydb. Brook Lobelia 272

74 Hieracium umbellatum L. Narrow-Leaved Hawk-weed 276

74 Agoseris aurantiaca (Hook.) Greene. Agoseris aurantiaca 276

75 Erigeron acris L. Blue Fleabane 288

75 Erigeron salsuginosus (Rich.) A. Gray. Large Purple Fleabane 288

75 Saussurea densa Hook. Saussurea 288

76 Erigeron discoideus Rydb. Cut-Leaved Fleabane 292

76 Erigeron melanocephalus A. Nelson. Black-Headed Fleabane 292

76 Erigeron multifidus Rydb. Cut-Leaved Fleabane 292

77 Aster Lindleyanus T. & G. Aster Lindleyanus 298

78 Erigeron aureus Greene. Golden Fleabane 302

78 Gaillardia aristata Pursh. Great-Flowered Gaillardia 302

79 Arnica cordifolia Hook. Heart-Leaved Arnica 304

79 Arnica louiseana Farr. Pale-Flowered Arnica 304

GLOSSARY

Acaulescent: with stem so short that leaves appear to arise

directly from root.

Achene: a small, dry, one-seeded indehiscent, seed-like fruit.

Acuminate: tapering gradually to a point.

Acute: with an abrupt point.

Adnate: growing together as the stamens with the petals,

or the anthers with the filament.

Alternate: (leaves) not opposite but arising at different points on the

stem. Stamens are alternate with petals when they are situated

in the intervals.

Anther: the part of the stamen which contains the pollen.

Apical: pertaining to the top or apex.

Auriculate: with ear-like appendages at base.

Axial: pertaining to an axis.

Bilabiate: two-lipped.

Blade: the expanded portion of a leaf.

Bract: a modified and usually small leaf at the base of a flower or

flower-cluster. In the latter case it is usually associated with others.

Bracteolate: furnished with small bracts.

Cæspitose: growing in tufts.

Calyx: the outer whorl of floral leaves forming usually a green cup.

Campanulate: bell-shaped.

Canescent: greyish-white, due usually to fine hairs.

Capillary: hair-like.

Capitate: arranged in a head; knob-like.

Capsule: a dry, dehiscent seed-vessel.

Carpel: a pistil-leaf. The pistil is regarded as being composed of

one or more such carpels.

Catkin: the scaly, cylindrical flower-cluster of Willow, Birch, etc.

Caudate: with a tail-like appendage.

Caudex: the perennial rootstock of an otherwise annual plant.

Chartaceous: papery or parchment-like.

Ciliate: fringed with marginal hairs.

Cinereous: ashen or greyish.

Cleistogamic: (flowers) never opening as the late flowers of

violets, etc.

Connate: united.

Cordate: heart-shaped.

Coriaceous: leathery.

Corm: a solid bulb.

Corymbose: with a flat-topped or convex flower-cluster,

the flower-stalks of which arise at different

points on the axis, and of which the outermost

flowers are the oldest.

Crenate: with rounded teeth.

Crenulate: with small rounded teeth.

Cuneate: wedge-shaped.

Cuspidate: ending in a sharp, hard point.

Cyme: a flat-topped or convex flower-cluster, the flower-stalks of

which arise at different points on the axis, and of which the

corolla innermost flowers are the oldest.

Deciduous: subject to fall, as leaves of most trees and calyx and

corolla of most flowers before the fruit forms.

Decumbent: (stems) reclining on the ground with summit

tending to rise.

Decurrent: (leaves) with base prolonged below insertion

and forming a sort of wing.

Deltoid: broadly triangular.

Dentate: toothed, usually with teeth projecting outwards.

Denticulate: with little, outwardly projecting teeth.

Diœcious: having staminate and pistillate flowers

on separate plants.

Disc: the central part of a head as opposed to the ray,

e. g., in the sun-flower. Also a fleshy expansion

of the receptacle of the flower.

Drupe: a stone-fruit.

Drupelet: a small stone-fruit.

Elliptic: with the outline of an ellipse.

Entire: with the margin even and not toothed, lobed, or divided.

Erose: irregularly margined.

Fimbriate: fringed.

Floccose: with loose tufts of wool-like hairs.

Foliaceous: leaf-like.

Follicle: a fruit of one carpel (pistil-leaf) which splits only

by one suture.

Fusiform: spindle-shaped.

Galea: a helmet-shaped body.

Glandular: with small prominences which appear to have

a secreting function.

Glaucous: covered with a bluish or white bloom.

Globose: spherical.

Incised: sharply and irregularly cut into.

Indusium: the membrane covering the fruit-dots of ferns.

Inferior: (side of flower) the one away from the axis of the

flower-cluster; (ovary) united with the calyx tube, the

other floral parts apparently growing from its summit.

Inflorescence: the mode of arrangement of flowers on the

stem; the flower-cluster.

Internode: the portion of the stem between the places where

the leaves arise.

Involucre: a whorl of small leaves at the base of a flower

or flower-cluster.

Keel: the lower petals of the peculiar corolla of the

pea, etc., which are united in the form of the keel of a boat.

Laciniate: with deep, triangular, narrow lobes.

Lanate: woolly.

Lanceolate: narrow and tapering at each end.

Lateral: pertaining to the side.

Leaflet: one of the divisions of the blade of a compound leaf.

Linear: narrow and flat with the margins parallel.

Lyrate: lyre-shaped.

Monœcious: having the separate staminate and pistillate flowers

on the same plant.

Mucronate: with a short, abrupt point.

Node: the point of the stem from which one or more leaves arise.

Oblanceolate: lance-shaped with the base tapering

downwards.

Obtuse: blunt.

Orbicular: circular.

Ovate: egg-shaped (applied to flat bodies).

Ovoid: egg-shaped (applied to solids).

Palmate: (compound leaf) with the divisions so arranged that

they radiate from the petiole-like the fingers of the hand.

Panicle: a branched, flower-cluster with the older

flowers outermost.

Paniculate: with much branched open flower clusters.

Papillæ: minute, blunt projections.

Pappus: the bristles, awns, or hairs which crown the fruit

of the Composites.

Parietal: (placentæ) attached to the wall of the ovary.

Peduncle: a flower-stalk.

Perennial: lasting from year to year.

Perfoliate: (leaves) having the basal part so extended

and united as to make it appear as though the stem

grew through them.

Perianth: the calyx and corolla collectively.

Petal: a corolla-leaf.

Petaloid: petal-like.

Petiole: the leaf-stalk.

Pinnæ: the primary divisions of a compound leaf.

Pinnate: compound with leaflets arranged along a common axis.

Pinnatifid: pinnately cleft.

Pinnules: secondary pinnæ.

Placentæ: the parts of the ovary to which the seeds are attached.

Plumose: feathery.

Pollen: the fertilising powder of the anther.

Polygamous: having both perfect and imperfect flowers.

Prostrate: lying flat on the ground.

Puberulent: covered with tiny almost imperceptible hairs.

Pulvini: cushions or pads.

Raceme: a flower-cluster in which the stalked flowers are

arranged along an elongated axis.

Racemose: raceme-like.

Rachis: the axis.

Radical: arising from the root, or portion of the stem

resembling the root.

Receptacle: the usually somewhat enlarged portion of the stem

on which parts of a flower or of a flower-cluster are borne.

Reniform: kidney-shaped.

Repand: wavy-margined.

Reticulate: arranged like a network.

Revolute: rolled backward.

Rhizome: an underground stem; a rootstock.

Rhombic: having the shape of a four-sided figure with parallel

and equal sides which may be either perpendicular or

oblique to each other.

Rhomboid: of a somewhat rhombic shape.

Rotate: wheel-shaped.

Runcinate: coarsely cut into with the segments turned backward.

Sagittate: shaped like an arrow-head with the lobes

turned downward.

Samara: a simple, indehiscent, winged fruit.

Saprophyte: a plant which grows on dead organic matter.

Scale: a minute, rudimentary leaf.

Scape: a leafless or nearly leafless flower-stalk arising

from near the ground.

Scarious: dry, thin, membranous.

Scurf: minute scales on the epidermis (skin).

Sepals: the divisions of the calyx.

Serrate: with teeth pointing forwards.

Serrulate: with minute forward pointing teeth.

Sessile: (leaf) without any stalk.

Sheath: a tubular structure such as is formed by the

lower part of the leaves of grasses, etc.

Silicle: a short and broad silique.

Silique: the elongated two-valved and usually dehiscent

fruit of the Cress family. It is divided

by a partition and the placentæ are parietal.

Sinuate: with a strongly wavy margin.

Sinus: the recess between the lobes of a leaf.

Sori: the fruit-dots of ferns.

Spadix: a fleshy spike as in the Araceæ.

Spathe: a large bract or pair of bracts enclosing a

flower or an inflorescence.

Spatulate: spoon-shaped.

Spike: a flower-cluster in which the flowers are stalkless

on an elongated axis.

Sporanges: the sacs containing the spores.

Spore: the reproductive body of ferns.

Sporophyll: leaf-like organ or receptacle containing spores.

Stamen: the organ of the flower which produces the pollen.

Staminodia: sterile stamens or structures which are developed

in the place of stamens.

Stellate: star-shaped.

Stipe: the leaf-stalk of a fern.

Stipule: an appendage such as is found at the base of many

leaf-stalks.

Stolon: a trailing and rooting shoot.

Striate: marked with slender, longitudinal stripes.

Strict: straight and upright.

Style: (of pistil) the elongated portion at top of ovary.

Subulate: awl-shaped.

Superior: (side of flower) the one next the axis of a flower-cluster;

(ovary) free from calyx tube.

Terete: cylindrical, circular in cross-section.

Ternate: in threes.

Tomentose: clothed with matted wool-like hairs.

Torulose: knobby; swollen at intervals.

Truncate: as if cut off at the top.

Tubercle: a small knob-like outgrowth.

Turbinate: top-shaped.

Umbel: an umbrella-like flower-cluster.

Undulate: wavy.

Valvate: (leaves in bud) meeting by the margins, not overlapping.

Verticillate: whorled.

Vexillum: the standard of the peculiar flower of pea, etc.

Villous: with long soft hairs.

GENERAL KEY TO THE FAMILIES

PLANTS WITHOUT FLOWERS

Ferns and their Allies

Leaves large, fern-like.

Sporanges in a spike or panicle above the sterile leaf. Ophioglossaceæ.

Sporanges borne in sori, on the backs or margins of the leaves Polypodiaceæ.

Leaves small, reduced to scales.

Stems erect, hollow, jointed, rush-like; leaves reduced to sheaths

at the joints; sporanges in the terminal, cone-like spike. Equisetaceæ.

Stems not hollow, creeping; leaves numerous, scale-like; sporanges

of one kind, solitary in the axils of the leaves. Lycopodiaceæ.

Stems creeping, moss-like; leaves numerous; sporanges of two kinds

in the axils of the upper leaves, which are formed into a

quadrangular spike. Selaginellaceæ.

PLANTS WITH FLOWERS

Woody Stemmed Plants; Trees, Shrubs, and Vines

TREES OR SHRUBS WITH RESINOUS JUICE; LEAVES USUALLY EVERGREEN.

Fruit a cone, either dry or berry-like. Pinaceæ.

Fruit a fleshy crimson berry. Taxaceæ.

TREES, SHRUBS, OR VINES WITH WATERY JUICE.

Leaves deciduous.

Flowers in catkins.

Staminate and pistillate catkins on different plants;

seed with white, silky hairs. Salicaceæ.

Staminate and pistillate catkins on the same plant;

fruit cone-like. Betulaceæ.

Flowers not in catkins.

(A) Trees.

Leaves simple, not lobed; flowers white. Drupaceæ.

Leaves lobed; flowers greenish-yellow. Aceraceæ.

Leaves pinnate; flowers white. Pomaceæ.

(B) Shrubs or vines.

*Stems spiny erect or trailing.

(a) Leaves simple or lobed.

Flowers small in dense terminal racemes. Araliaceæ.

Flowers few in axillary clusters or racemes. Grossulariaceæ.

(b) Leaves pinnate or palmate. Rosaceæ.

**Stems not spiny but with minute scales;

fruit silver, scarlet or orange berries. Elæagnaceæ.

***Stems erect, smooth.

(a) Petals distinct.

†Leaves simple.

Flowers small, in flat heads or clusters.

Petals 4; stamens 4. Cornaceæ.

Petals 5; stamens numerous. Rosaceæ.

Flowers in axillary clusters or racemes; petals 5; stamens numerous.

Fruit a berry. Drupaceæ.

Fruit a pome. Pomaceæ.

††Leaves pinnate; flowers yellow. Rosaceæ.

(b) Petals united.

Flowers small, short-campanulate in axillary

or terminal heads. Caprifoliaceæ.

Flowers large, funnel-form, in terminal

or axillary clusters. (Azaleastrum) Ericaceæ.

Flowers bell or urn-shaped, few, axillary Vacciniaceæ.

****Stems creeping or twining, smooth or hairy.

Flowers with distinct petals, blue or purple. (Atragene). Ranunculaceæ.

Flowers with united petals.

Corolla tubular. Caprifoliaceæ.

Corolla bell-shaped. Ericaceæ.

Leaves evergreen; shrubs or vines.

Leaves pinnate; flowers yellow. Berberidaceæ.

Leaves simple.

Flowers inconspicuous; fruit a black berry. Empetraceæ.

Flowers showy, open, campanulate or urn-shaped,

white or pink. Ericaceæ.

Herbaceous Plants

Plants without leaves.

Parasitic on branches or stems of evergreens. Loranthaceæ.

Not parasitic on trees; roots coral-like. (Corallorhiza) Orchidaceæ.

Plants with leaves.

Leaves parallel-veined.

(A) Flowers small, collected on a spadix. Araceæ.

(B) Flowers not collected on a spadix.

*Flowers regular; corolla lobes 6; stamens 6.

(a) Leaves grass-like, mostly at the base.

Flowers white or greenish. Melanthaceæ.

Flowers pink. (Allium). Liliaceæ.

Flowers blue. (Sisyrinchium). Iridaceæ.

(b) Leaves not grass-like, mostly on the stem.

Flowers white or pink. Convallariaceæ.

Flowers green. Melanthaceæ.

Flowers yellow or orange. Liliaceæ.

Flowers red or wine-color. (Kruhsea.) Convallariaceæ.

**Flowers irregular, frequently with broad or inflated lip and spur. Orchidaceæ.

Leaves not parallel-veined.

(A) Flowers small, in close heads surrounded by an involucre.

Flowers all alike, florets ligulate. Cichoriaceæ.

Flowers not alike, central ones tubular, marginal usually with rays. Carduaceæ.

(B) Flowers in umbels at the ends of the stems or branches.

Umbels broad, flat or nearly so, flowers white, yellow, or purple. Apiaceæ.

Umbels round.

Flowers greenish; leaves compound. Araliaceæ.

Flowers white or pink; leaves simple. (Eriogonum.) Polygonaceæ.

(C) Flowers not in umbels.

*Flowers regular.

(a) Flowers with 2 sepals and 5 petals; corolla pink. Portulacaceæ.

(b) Flowers with 3 sepals and 3 petals; seeds triangular. Polygonaceæ.

(c) Flowers with 4 sepals and 4 petals.

Stem angular; leaves in whorls. Rubiaceæ.

Stem round; leaves not in whorls

Stamens 6, erect. Brassicaceæ.

Stamens 8, spreading. Onagraceæ.

(d) Flowers with 5 sepals or petals, distinct.

†Stamens as many or twice as many as the corolla lobes.

Stems swollen at the nodes. Caryophyllaceæ.

Stems not swollen at the nodes

Stem leaves numerous, entire, thin; flowers in

terminal or axillary clusters, white or greenish. Santalaceæ.

Leaves basal, entire; flowers solitary on a 1-leaved scape. Parnassiaceæ.

Leaves mostly basal, lobed or cut; flowers in

a close spike or head. Saxifragaceæ.

Leaves and stems fleshy.

Follicles 2. Saxifragaceæ.

Follicles 4. Crassulaceæ.

Leaves numerous, stiff, awl-shaped; stems low, tufted. Saxifragaceæ.

††Stamens more than twice as many as the corolla lobes

§Leaves basal or mostly so.

Leaves evergreen, simple or lobed, flowers yellow

or white, solitary on naked scapes. (Dryas.) Rosaceæ.

Leaves not evergreen, simple, lobed, or much divided

Smooth.

Stems erect or floating. Ranunculaceæ.

Stems creeping.

Flowers white or pink. Rosaceæ.

Flowers yellow. Ranunculaceæ.

Rough-hairy.

Flowers yellow or pink. Rosaceæ.

Flowers white, blue or purple. Ranunculaceæ.

§§Leaves mostly on the stem.

Leaves finely cut; stems creeping; flowers white. Rosaceæ.

Leaves entire.

Leaves bright green, black-dotted; flowers yellow. Hypericaceæ.

Leaves glaucous, without black dots; flowers blue. Linaceæ.

(e) Flowers with 5 sepals and 5 united petals or lobes.

†Stamens 3; flowers small, in terminal clusters. Valerianaceæ.

††Stamens 5.

Opposite to the corolla lobes. Primulaceæ.

Alternate with the corolla lobes.

Stems smooth, leafy; corolla campanulate.

Stem leaves broad; flowers pink. Apocynaceæ.

Stem leaves narrow; flowers blue. Campanulaceæ.

Stems rough-hairy.

Corolla salver-form; stamens included. Boraginaceæ.

Corolla tubular; stamens exserted. Hydrophyllacæ.

†††Stamens 10; flowers mostly in terminal heads or singly. Pyrolaceæ.

**Flowers irregular.

(a) Leaves all entire and basal, forming rosettes; flowers blue. Lentibulariaceæ.

(b) Leaves all heart-shaped; flowers blue, white or yellow. Violaceæ.

(c) Leaves not heart-shaped, simple or much cut, not pinnate.

Stems square; flowers in close verticels in the axils of

the leaves or in terminal heads. Menthaceæ.

Stems round; flowers numerous in elongated terminal leafy heads

or few on slender stems in the axils of the upper leaves. Scrophulariaceæ.

(d) Leaves palmate; flowers blue in elongated spikes. Ranunculaceæ.

(e) Leaves pinnately divided; flowers pea-like. Papilionaceæ.

(f) Leaves compoundly lobed, not pinnate.

Stems low, spreading, flowers small in dense terminal.

or axillary spikes. Papaveraceæ.

Stems erect; flowers drooping in open terminal heads, either showy and spurred

or not showy and without spurs; leaves maiden-hair fern-like. Ranunculaceæ.

Alpine Flora of the Canadian Rocky Mountains

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