Note 111 — PART III.—THE METAMORPHOSES OF INSECTS (continued 2)

In his remarkable studies on the morphology of the Lepidoptera, Professor W. Hatchett-Jackson states his belief that Riley’s homology of the sustentors with the soles or plantæ of the anal prolegs, and the sustentor ridges with their limbs, is wrong, and that the eminences on either side the anal furrow, or the “anal prominences,” as they are termed by Riley, represent the prolegs, and that the sustentor ridges and sustentors are probably peculiar developments of the body of the 10th somite, found only in some Lepidoptera. From our examination of pupa of different families of moths, we are satisfied that Jackson’s view is the correct one. We have not found the sustentors and their ridges in the pupæ of the more generalized moths, but the vestiges of the anal legs are almost invariably present, their absence in the pupa of Nola and Harrisina being noteworthy.