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The Vertebrate Skeleton

Note 173 — Chapter XXIII · GENERAL ACCOUNT OF THE SKELETON IN MAMMALIA (continued).

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See E. Cope, "The origin of the foot structures of Ungulata," Journ. of Philad. Acad. 1874. H.F. Osborn, "The evolution of the Ungulate foot," T. Amer. Phil. Soc. 1889.

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