Note 35 — Chapter VII · GENERAL DIFFERENCES OF THE TWO LIVES WITH RESPECT TO VITAL POWER.
F. Gold translation
The motions of the iris cannot be attributed to an active expansion of an erectile texture; they are owing to the contractions of two muscular layers, one of which is radiated and enlarges the opening of the pupil, the other is orbicular and contracts it.
The motions of the iris, like all those which have muscular contraction for their cause, can be excited for a considerable time after death by the galvanic fluid. During life, the motions of the pupil are produced in man, by the more or less vivid impression of light on the retina. But they are beyond the influence of the will; in birds on the contrary, they appear to be entirely subjected to it. In these animals, we can even after death, and on an eye entirely detached from the body, produce the motions of the iris by pricking the optic nerve.