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On the Variation of Species, with Especial Reference to the Insecta

Note 64 — Chapter V · GEOLOGICAL REFLECTIONS.

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Assuming the Helix Lowei and Bowdichiana to be gigantic phases of the H. Portosanctana and punctulata, respectively; four only, namely H. fluctuosa and lapicida, Achatina Eulina, and Cyclostoma lucidum (the first three of which are extinct throughout the entire group), seem to have altogether disappeared. Nevertheless, the gradual dying-out, as it were, of species, both here and in Madeira proper, is singularly evident. Thus, in the latter, the Caniçal beds show the H. tiarella to have been once most abundant (it literally teems in those calcareous formations). Yet so rare is it in a recent state, that, until the summer of 1855, when it was detected by myself and the Rev. R. T. Lowe in two remote spots along the perpendicular cliffs of the northern coast, it was supposed to have been lost for ages. And the same may be said of its counterpart, the H. coronata, in Porto Santo,—which, likewise, swarms in every fossil-bed of that island; but which was, also, until I met with it, on the 15th of December 1848, adhering to slabs of stone at a considerable depth beneath the ground, on the extreme eastern peak (opposite to the Ilheo de Cima), imagined to have long passed away. And so, reasoning from analogy, I think it far from improbable that the third representative of this little geographical assemblage,—the H. coronula of the Bugio (which has hitherto only occurred in the mud deposits on the summit of that rock),—may be still alive, though perhaps in very small numbers, on some of the inaccessible ridges of those dangerous heights.

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