Note 7 — Chapter IX
Reading edition
Other similar cases of factorial repulsion have been demonstrated in the sweet pea, and two of these are also concerned with the two factors with which we have just been dealing. Two distinct varieties of pollen grains occur in this species, viz. the ordinary oblong form and a rather smaller rounded grain. The former is dominant to the latter.[7] When a cross is made between a purple with round pollen and a red with long pollen the F1 plant is a long pollened purple. But the F2 generation consists of purples with round pollen, purples with long pollen, and reds with long pollen in the ratio 1 : 2 : 1. No red with round pollen appears in F2 owing to repulsion between the factors for purple (B) and for long pollen (L). Similarly plants produced by crossing a red hooded long with a red round having an erect standard give in F1 long pollened reds with an erect standard, and these in F2 produce the three types, round pollened erect, long pollened erect, and long pollened hooded, in the ratio 1 : 2 : 1. The repulsion here is between the long pollen factor (L) and the factor for the erect standard (E).